Revision of Key to Genera from Wed, 2011-05-18 23:28

1.









Culms woody; leaves pseudopetiolate

2

 

Culms usually herbaceous; if woody the leaves not pseudopetiolate

5

 
2.

Culms forming climbing vines

Mullerochloa

 

Culms not forming climbing vines

3

 
3.

Auricles with bristles to 10 mm long

Neoleleba

 

Auricles without bristles

4

 

4.

Culms cylindrical in cross section

Bambusa

Culms with an internodal groove on alternate sides

*Phyllostachys

5.

Leaves spirally arranged

Micraira

Leaves arranged in two rows

6

6.

Spikelets morphologically or functionally unisexual

7

Spikelets bisexual

20

7.

Spikelets with 1 floret, or more than 2 florets

8

Spikelets with 2 florets

12

8.

Spikelets with 1 floret; male and female spikelets on same plant

9

Spikelets with more than 2 florets; male and female spikelets on different plants

10

9.

Lemma asymmetrical with a ±lateral orifice

Leptaspis

Lemma pyriform with a terminal orifice

Scrotochloa

10.


11.

Short grasses with a compact raceme

 Distichlis

Tall reed-like grasses with a plumose panicle

11

Leaf blades with only the midrib prominent and strongly sclerified

*Cortaderia

 Leaf blades with several wide strongly sclerified veins (including midrib)

*Austroderia        

12.

Upper lemma membranous, thinner than glumes

13

Upper lemma hardened, thicker than glumes

16

13.

Male and female spikelets in separate inflorescences

*Zea

Male and female spikelets in different parts of the same inflorescence

14

14.



15.


Inflorescence consisting of a female spikelet enclosed in a rigid shell, from which a raceme of male spikelets emerges apically

Coix

Inflorescence spike-like with female spikelets basal and male apical

15

Internode broader than and ± enclosing female spikelet

Chionachne

Internode narrower  than and ± enclosed by female spikelet

*Tripsacum

16.

Plants monoecioeus, i.e. with male and female spikelets on the same plant

17

Plants dioecious i.e. with male and female spikelets on different plants

18

17.

Inflorescence a simple raceme, with 1-2 female spikelets at the base, the upper spikelets male

Thuarea

Inflorescence a spike-like panicle with racemes of female spikelets below and solitary male spikelets above

Hygrochloa

18.

Inflorescence a contracted, spike-like panicle, not spatheate

Pseudochaetochloa

Inflorescences clustered together in dense, umbel-like spatheate heads

19

19.

Female heads with long stiff spines; inhabits coastal dunes

Spinifex

Female inflorescences bracteate but not spiny; inhabits desert sand dunes

Zygochloa

20.

Spikelets with 1 to many florets, if 2-flowered both florets or lower one bisexual

21

Spikelets 2-flowered, lower floret male or barren, upper bisexual or female

167

21.

Inflorescence a solitary bilateral spike or raceme; spikelets usually with the broad side facing the axis

22

Inflorescence a panicle or a unilateral or multilateral spike or raceme, rarely a bilateral spike but then the spikelets edgewise on to the axis

38

22.

Lemmas 3-5-nerved, hyaline to membranous

23

Lemmas 5-10-nerved, hardened at maturity

26

23.

Lemmas awned

*Psilurus

Lemmas not awned

24

24.

Spikelets compressed laterally

*Parapholis

Spikelets compressed dorsiventrally

25

25.

Lemmas with lateral nerves extending to the apex

Lepturus

Lemmas with very short lateral nerves

*Hainardia

26.

Rachilla terminated by a fertile floret

Tetrarrhena

Rachilla prolonged beyond the uppermost fertile floret

27

27.

Spikelets in groups of more than 2 at each rachis node

28

Spikelets solitary at each rachis node

26

28.

Spikelet 4-5-flowered, in groups of 2-4 (-7)

*Leymus

Spikelet 1-flowered, in groups of 2 or 3

30

29.

Spikelets in pairs at each rachis node

*Taeniatherum

Spikelets in 3s at each rachis node

*Hordeum

30.

Annuals

31

Perennials

34

31.

Glumes not keeled

Brachypodium

Glumes strongly keeled

32

32.

Glumes subulate

*Secale

Glumes not subulate

33

33.

Lemmas with nerves converging upwards

*Eremopyrum

Lemmas with nerves not converging upwards

*Triticum

34.

Rachis fragile, smooth

*Thinopyrum

Rachis tough, scabrous

35

35.

Plant with long creeping rhizomes

*Elytrigia

Plant tufted, or with short rhizomes

36

36

Leaf blades ± flat; glumes ± scabrous on nerves

Anthosachne

Leaf blades usually convolute or involute; glumes smooth

37

37.

Rachis internodes concave; glumes subacute to pointed

Australopyrum

Rachis internodes nearly flat; glumes truncate

*Thinopyrum

38.

Fertile lemmas deeply or distinctly lobed

39

Fertile lemmas entire, dentate or slightly lobed

42

39.

Spikelets 1-flowered

Amphipogon

Spikelets 2-or more-flowered, rarely 1-flowered

40

40.

Lemmas 9-lobed

Enneapogon

Lemmas 3-lobed

41

41.

Inflorescence a 1-sided raceme

Astrebla

Inflorescence a panicle or spike

Triodia

42.

Leaf blades hard, woody, needle-like

Triodia

Leaf blades not hard, woody, needle-like

43

43.

Leaf blades with transverse veins between main veins

44

Leaf blades without transverse veins

45

44.

Glumes more or less equal; upper glume 7-nerved; lemmas awned

Lophatherum

Glumes very unequal; upper glume 3-5-nerved; lemmas mucronate

Centotheca

45.

Tall reed-like plants

46

Plants not tall and reed-like

47

46.

Lemmas hairy; rachilla not hairy

*Arundo

Lemmas not hairy; rachilla hairy

Phragmites

47.

Spikelets with 2 or more bisexual florets or if only one, with rachilla extension or sterile florets (empty lemmas) above it

48

Spikelets with 1 bisexual floret, with or without male or sterile florets (empty lemmas) below it

129

48.

Spikelets with 3 or more florets or with 1or 2 florets and a rachilla extended beyond the upper floret

49

Spikelets with 2 florets, without a rachilla extension

122

49.

Inflorescence racemose or spicate, or digitate or non-digitate

50

Inflorescence a panicle

76

50.

Inflorescence digitate or subdigitate

51

Inflorescence racemose or spicate

62

51

Spikelet with at least 3 fertile florets

52

Spikelet with 1 (rarely 2) fertile florets

55

52.

Axis of each inflorescence branch ending in a bristle; spikelet-bearing axis disarticulating

Dactyloctenium

Axis of each inflorescence branch ending in a spikelet; spikelet-bearing axis not disarticulating

53

53.

Spikelets 2-5; lemma 5-7-nerved; leaves mostly basal, setaceous

Cyperochloa

Spikelets many; lemma 3-nerved; leaves not basally aggregated or setaceous

54

54.

Lemmas mucronate to shortly awned; upper glume 1-nerved

Acrachne

Lemmas muticous; upper glumes 3-7-nerved

*Eleusine

55.

Spikelet with 1 floret; lemmas unawned

56

Spikelet with 2 or more florets; lemmas awned

57

56.

Glumes shorter than floret

Cynodon

Glumes longer than floret

Brachyachne

57.

Lowest lemma dorsally compressed (lying on front or back when placed on a flat surface)

58

Lowest lemma laterally compressed (lying on the side when placed on a flat surface)

59

58.

Glumes unequal, shorter than florets

Enteropogon

Glumes subequal, longer than florets

Austrochloris

59.

Inflorescence with more than 40 branches

*Eustachys

Inflorescence with less than 20 branches

60

60.

Awn of lowest lemma less than 2 mm long

x Cynochloris

Awn of lowest lemma more than 3 mm long

61

61.

Lemmas very broad, wing-like

Oxychloris

Lemmas not very broad, not wing-like

Chloris

62.

Spikelets unilateral (borne on one side of the inflorescence axis)

63

Spikelets bilateral or multilateral (borne on opposite sides or all round the inflorescence axis)

70

63.

Spikelet with many fertile florets

64

Spikelet with one fertile floret

66

64.

Lower glume 5-nerved; lemma 7-nerved

Tribolium

Lower glume 1-nerved; lemma 1-3-nerved

65

65.

Lemma mucronate to shortly awned; spikelet rachilla and callus not hairy

Tripogon

Lemma awnless; spikelet rachilla and callus hairy

Eragrostiella

66.

One glume only developed

67

Both glumes well developed

68

67.

Spikelet with one floret; lemma 3-nerved

*Nardus

Spikelet with many florets; lemma 3-nerved

*Lolium

68.

Spikelets falling entire; spikelets 6-18 mm long

*Spartina

Spikelets fragmenting above glumes; spikelets less than 5.5 mm long

69

69.

Spikelets shortly pedicellate

*Mibora

Spikelets sessile

Microchloa

70.

Spikelets disarticulating above glumes; rachilla extending beyond upper floret

71

Spikelets falling with glumes; rachilla terminated by a floret

73

71.

Spikelets sessile, partially embedded in the rachis

*Gaudinia

Spikelets pedicelled, not embedded in the rachis

72

72.

Lemmas awned; glumes not keeled

*Vulpia

Lemmas awnless; glumes keeled

*Catapodium

73.

Inflorescence a true spike; spikelets sessile, sunken in hollows along the axis

*Pholiurus

Inflorescence a spike-like raceme or spike-like panicle; spikelets at least shortly pedicellate and not sunken in hollows

74

74.

Spikelets burr-like with hooked spines

Tragus

Spikelets not burr-like

75

75.

Glume 1

Zoysia

Glumes 2

Perotis

76.

Glumes usually shorter than lowest floret and with the upper florets distinctly exserted; lemmas awnless or with a straight or curved awn from an entire or bifid apex, or several-awned or lobed

77

Glumes (or the longer one) at least as long as or longer than the lowest floret, often as long as the spikelet and enclosing the florets; lemmas awnless or more often awned from the back or from the sinus of a 2-lobed tip, the awn usually geniculate, rarely the awn straight or curved from the tip

103

77.

Lemmas 1-3-nerved

78

Lemmas 5-many nerved

87

78.

Lemmas 3-awned

Triraphis

Lemmas 1-awned or without awns

79

79.

Lemmas notched, toothed or lobed

80

Lemmas entire

82

80.

Glumes longer than lemmas, awned

*Dinebra

Glumes shorter than or equal to lemmas, not awned

81

81.

Spikelets borne in leaf-sheath axils

Psammagrostis

Spikelets not borne in leaf-sheath axils

Leptochloa

82.

Lemmas awned

83

Lemmas not awned

84

83.

Spikelets in globose clusters; palea winged

Elytrophorus

Spikelets in dense to loose panicles; palea not winged

Ectrosia

84.

Lemmas with winged keels

Heterachne

Lemmas without winged keels

85

85.

Ligule membranous

*Sphenopus

Ligule a fringe of hairs

86

86.

Lemmas 1-nerved

Thellungia

Lemmas 3-nerved

Eragrostis

87.

Ovary hairy at apex

88

Ovary glabrous at apex

90

88.

Leaf sheath margins joined

*Bromus

Leaf sheath margins free

89

89.

Leaf blades 7-24 mm wide; lemmas bilobed

Dryopoa

Leaf blades 2-3 mm wide; lemmas 3-lobed

Notochloë

90.

Spikelets dimorphous, with bisexual spikelets surrounded by sterile ones

91

Spikelets not dimorphous

92

91.

Spikelets disarticulating above the glumes; spikelet-bearing axes persistent

*Cynosurus

Spikelets disarticulating with the glumes; spikelet bearing axes disarticulating

*Lamarckia

92.

Inflorescence a contracted panicle or a panicle of racemes

93

Inflorescence an open panicle

97

93.

Glumes more or less equal

94

Glumes very unequal

96

94.

Glumes not keeled

*Melica

Glumes keeled

95

95.

Leaf blades up to 3 mm long and basal, if present

Spartochloa

Leaf blades well developed and not mainly basal

*Catapodium

96.

Glumes very dissimilar; lemmas awned

*Vulpia

Glumes similar; lemmas awnless

*Sclerochloa

97.

Spikelets almost as wide as long

*Briza

Spikelets up to half as wide as long

98

98.

Lemmas ± rounded or flattened on back, at least below

99

Lemmas keeled throughout

100


   

99.

Lemmas 7-9-nerved; leaf
sheath margins joined

Glyceria

Lemmas 5-nerved; leaf
sheath margins free

Puccinellia

100.

Leaves not basally
aggregated; lemmas firmer than glumes, awned

101

Leaves mostly basal;
lemmas similar in texture to glumes, awnless

102

101.

Lemmas keeled; palea
thinner than lemma

*Dactylis

Lemmas not keeled;
palea with similar texture to lemma

*Festuca

102

Glumes as long the
spikelet or lemmas mucronate; relative to the adjacent lemmas; rachilla hairy

Hookerochloa

Glumes much shorter
than the adjacent lemmas; lemmas muticous; rachilla hairless

Poa

103.

Ligule membranous

104

Ligule a fringe of
hairs, ciliate membrane or ciliate at margins

113

104.

Lemmas 3-nerved

*Avellinia

Lemmas 5-9-nerved

105

105.

Lower floret male

*Arrhenatherum

Lower floret bisexual

106

106

Glumes more than 14 mm
long

*Avena

Glumes less than 10 mm
long

107

107.

Annuals

108

Perennials

110

108.

109.

Panicle spicate

 *Rostraria

Panicle open

 109

Awn witha ring of
hairs at junction of column and limb

*Corynephorus

Awn without a ring of
hairs

*Aira

110.

Lemmas 4-pointed, or
2-lobed with each lobe 2-pointed

111

Lemmas entire,
2-pointed or 2-lobed but not as above

112

111.

Glumes long relative
to the adjacent lemmas; leaves mostly basal

Deschampsia

Glumes shorter than
the adjacent lemmas; leaves not basally aggregated

Amphibromus

112

Lemma awn 4-9 mm long,
geniculate

Trisetum

Lemma awnless or with
a short awn to 0.7 mm long

*Koeleria

113.

Leaf sheaths of the
lower leaves swollen and woolly or leaf blades hard, woody, needle-like

114

Leaf sheaths of the
lower leaves not swollen and woolly and leaf blades not hard, woody,
needle-like

115

114.

Spikelets with 3-8
fertile florets; leaf sheaths of the lower leaves swollen and woolly

Monachather

Spikelets with 1
fertile floret; leaf blades hard, woody, needle-like

Symplectrodia

115.

Lemmas awnless,
sometimes mucronate

116

Lemmas distinctly
awned

120

116.

Lemmas entire, if
mucronate, mucros apical

*Tribolium

Lemmas bilobed; if
mucronate, mucro is from an apical sinus

117

117.

Grain not
longitudinally grooved

*Schismus

Grain longitudinally
grooved

118

118.

Lemma only hairy on
the lower margins

*Danthonia

Lemma ± evenly
pubescent or in tufts

119

119.

Lemma ± evenly
pubescent in lower half only/td>

Plinthanthesis

Lemma with hairs over
whole surface or in tufts

Rytidosperma

120

Hilum ± 1/2 length of
grain

*Chionochloa

Hilum up to 1/3 length
of grain

121

121.

Lemma ± evenly
pubescent in lower half only

Plinthanthesis

Lemma with hairs over
whole surface or in tufts

122

122

Hairs on the lemma in
two untidy rows, most of lemma surface glabrous and shiny; leaves often disarticulating
from sheath

Rytidosperma

Hairs on the lemma
forming a continuous indumentum, sometimes with tufts on the upper margin;
leaves not disarticulating

123

123.

Lemmas either
distinctly hairy or awned

124

Lemmas neither
distinctly hairy nor awned

128

124.

Upper glume twice
length of lower glume

Pheidochloa

Glumes ± equal in
length

125

125.

Glumes (3-)5-11-nerved

Eriachne

Glumes 1-3-nerved

126

126.

Bisexual florets 1 per
spikelet

*Holcus

Bisexual florets 2 per
spikelet

127

127.

Lemma awn dorsal

Aira

Lemma awn terminal

*Pentameris

128.

Both lemmas
membranous, not hardened

*Periballia

Lower lemma hardened

Coelachne

129.

Glumes absent or
rudimentary

130

Glumes well developed

132

130.

Spikelets 1-flowered

Leersia

Spikelets 3-flowered,
the lower two reduced to sterile lemmas

131

131.

Lemma and palea
leathery to indurated

Oryza

Lemma and palea
membranous

Potamophila

132.

Spikelets with 3
florets, the lower two reduced to sterile or male and sometime minute lemmas

133

Spikelets with 1
floret

137

133.

Both sterile lemma
with a hinge-like appendage

*Ehrharta

Neither sterile lemmas
with an appendage

134

134.

Basal florets male

Hierochloë

Basal florets sterile

135

135.

Glumes minute

Microlaena

Glumes relatively
large

136

136.

Sterile lemmas shorter
than fertile floret; glumes winged

*Phalaris

Sterile lemmas longer
than fertile floret; glumes not winged

*Anthoxanthum

137.

Lemmas terete or
dorsally compressed

138

Lemmas laterally
compressed

145

138.

139.

Lemma with a
3-branched awn (very rarely 1-branched)

139

Lemma with a single
undivided awn

140

Awns glabrous

Aristida

Awns plumose

 *Stipagrostis

140.

Awn ± geniculate;
spikelet base (callus) pointed, often long and pungent

141

Awn ± straight;
spikelet base (callus) short and blunt

144

141.

Lemma margins usually
enclosing the palea; callus pungent; awn well developed and with a
strongly-twisted column at maturity

142

Lemma margins not
completely enclosing the palea; callus blunt; awns not as robust as above and
the column only weakly developed at maturity

143

142.

Hummock-forming
perennial; awn almost circular in transverse section

Monodia

Annuals or perennials,
not usually hummock-forming; awn flattened in transverse section

Austrostipa

143.

Lemma apex and lower
part of awn with hairs obscuring the base of the awn

Jarava

Lemma apex glabrous or
slightly hairy, the hairs not obscuring the base of the awn

Amelichloa

144.

Lemma smooth, with a
deciduous awn

*Piptatherum

Lemma strongly
scabrous, with a more or less persistent awn

*Nassella

145.

Lemmas coriaceous to
indurated

146

Lemmas membranous to
cartilaginous

151

146

Lemma awns
non-geniculate

147

Lemma awns geniculate

148

147.

Lemma with a
3-branched awn (very rarely 1-branched); spikelets with one floret

Aristida

Lemma with a single
awn; spikelets with 2 florets (very rarely 1)

Eriachne

148.

Lemma 1 awned and with
an apical crown

149

Lemma 3-5 awned and
without an apical crown

150

149.

Awn at least 15 mm
long; palea shorter than lemma, the lemma margins enclosing the palea

*Nassella

Awn less than 10 mm
long; palea slightly longer than lemma, the lemma margins exposing the palea

*Piptochaetium

150.

Lemma with 5 awns

Pentapogon

Lemma with 3 awns

Anisopogon

151.

Glumes similar in
texture to and shorter than lemmas; ligule hairy to some degree

152

Glumes firmer in
texture to and as long as or longer than lemmas; ligule an unfringed membrane

153

152.

Panicle spike-like,
subtended by an inflated sheath

*Crypsis

Panicle open to
contracted, without an inflated sheath

Sporobolus

153.

Spikelets falling
entire

154

Spikelets
disarticulating above the glumes

156

154.

Glumes blunt, ±united

*Alopecurus

Glumes acute or awned,
distinctly separate

155

155

Lemma 3-nerved

Garnotia

Lemma 5-nerved

Polypogon

156.

Glumes swollen at base

*Gastridium

Glumes not swollen at
base

157

157.

Glumes silky-woolly,
much longer than lemma

*Lagurus

Glumes not
silky-woolly, equal to or slightly longer than lemmas

158

158.

Spikelets at least 9mm
long

*Ammophila

Spikelets up to 7mm
long

159

159.

Glumes terminating
abruptly, shortly awned

160

Glumes obtuse or
acute, not or scarcely awned

161

160.

Lemmas awned;
inflorescence a contracted panicle

*x Agropogon

Lemmas not awned;
inflorescence a tight spike-like panicle

*Phleum

161.

Glumes with regularly
arranged hairs on keel

Echinopogon

Glumes smooth to rough
on keel, without hairs

162

162.

163.

Lemma awns more than
twice as long as lemma

163

Lemma awns, if
present, less than twice as long as lemma

164

Annuals

*Apera

Perennials

Dichelachne

164.

Lemmas decidedly
firmer than the glumes

Deyeuxia

Lemmas less firm than
the glumes

165

165.

Glumes at least 5.5mm
long

*Calamagrostis

Glumes less than 5mm
long

166

166.

Inflorescence axis
persistent; rachilla extension less than 20% of lemma length

Agrostis

Inflorescence often
detaching as a diaspore; rachilla extension 30-100% of lemma length

Lachnagrostis

167.

Spikelets breaking up
above the glumes

168

Spikelets falling
entire, breaking below the glumes

169

168.

Glumes very unequal;
spikelets compressed laterally

Arundinella

Glumes ±equal;
spikelets compressed dorsiventrally

Isachne

169.




170.


Spikelets often
dissimilar and usually paired, with one sessile and the other pedicelled
(occasionally both pedicelled, but unequally, or in 3's with 2 sessile and 1
pedicelled) or rarely with the pedicelled spikelet reduced to the pedicel;
upper lemma of sessile spikelet usually geniculately awned; glumes firm in
texture

170

Spikelets all alike
and usually solitary, rarely paired; upper lemma not geniculately awned;
glumes usually membranous

206

 Spikelets similar in shape and size, development
of awns and sex; if different pedicelled spikelets awnedand sessile unawned;
pedicels and raceme joints slender

171

Spiklets different in
shape and size, development of awns and sex; if similar, pedicels and
joints thickened

179

171.

Paired spikelets
unequally pedicelled

172

Paired spikelets with
at least one

sessile

174

172.

Spikelets awnless

Imperata

Spikelets awned from
apex of the upper

lemma

173

173.

Inflorescence a
panicle, the racemes at

least 10 cm long

*Miscanthus

Inflorescence of
subdigitate racemes up to

7 cm long

Pseudopogonatherum

174.

Inflorescence a much
branched panicle

Saccharum

Inflorescence a
solitary raceme, or a

digitate or subdigitate cluster of racemes

175

175.

Spikelets in 3s, with
2 sessile and 1

pedicelled

Polytrias

Spikelets in pairs (3s
at apex), with 1

sessile and 1 pedicelled

176

176.

Sessile spikelets
male; pedicelled

spikelets female

Germania

Sessile and pedicelled
spikelets of the

same sex

177

177.

Lower glume convex on
back, not

2-keeled

Pogonatherum

Lower glume flat or
grooved on back,

2?keeled

178

178.

Tufted grass with
silky hairs on

racemes

Eulalia

Creeping grass with no
hairs on

racemes

Microstegium

179.

Rachis joints and
pedicels thick, ±

contiguous or fused together, forming cavities into which the spikelets are

sunk

180

Rachis joints and
pedicels slender, or if

thickened upwards, not forming cavities

188

180.

Sessile spikelet awned

181

Sessile spikelet
awnless

183

181.

Pedicelled spikelets
reduced to naked

pedicels

Thelepogon

Pedicelled spikelets
developed

182

182.

Pedicels ± flat

Sehima

Pedicels swollen

Ischaemum

183.

Lower glume of sessile
spikelet

2-toothed

Elionurus

Lower glume of sessile
spikelet not

2-toothed

184

184.

Spikelets in opposite
rows on axis

Ophiuros

Spikelets on one side
of axis

185

185.

Spikelets of each pair
similar, both

bisexual; rachis not fragmenting

Hemarthria

Spikelets of each pair
dissimilar,

pedicelled male or sterile or spikelets borne singly; rachis fragmenting

readily

186

186.

Lower floret of
sessile spikelet male

Rottboellia

Lower floret of
sessile spikelet

sterile

187

187.

Lower glume of sessile
spikelet with a row

of marginal spines

Eremochloa

Lower glume of sessile
spikelet without

marginal spines

Mnesithea

188.

Inflorescence with
spathes, spatheoles or

spathe-like sheaths enclosing or subtending inflorescence

189

Inflorescences
espatheate

197

189.

Raceme 1 jointed,
reduced to 3

spikelets

190

Racemes with several
joints, with more than

3 spikelets

191

190.

Racemes in tight
clusters; lemmas mucronate

or shortly awned

Apluda

Racemes loose and
pedunculate; lemmas with

long awns

Clausospicula

191.

Fertile spikelets
±compressed, the lower

glume distinctly 2-keeled for most of its length

192

Fertile spikelets
±terete, the lower glume

2-keeled only at the tip if at all

195

192.

Lower part of awn as
wide as the lemma from

which it arises

Spathia

Lower part of awn
narrower than the lemma

from which it arises

193

193.

Raceme bases terete

*Andropogon

Raceme bases flattened

194

194.

Racemes paired; leaves
aromatic

Cymbopogon

Racemes solitary;
leaves not aromatic

Schizachyrium

195.

Fertile spikelets not
within an involucre

of male or barren spikelets

Hyparrhenia

Fertile spikelets
within an involucre of

male or barren spikelets

196

196.

Fertile spikelets
falling with involucre or

whole raceme falling as one unit; callus blunt

Iseilema

Fertile spikelets
falling separately from

involucres; callus pungent

Themeda

197.

Rachis joints and
pedicels with a

translucent mid-line between thickened margins

198

Rachis joints and
pedicels without a

translucent mid-line

199

198.

Inflorescence more
than once-branched

Capillipedium

Inflorescence an
arrangement of racemes on

a central axis

Bothriochloa

199.

Inflorescence of
solitary, paired or

subdigitate racemes

200

Inflorescence a
panicle

202

200.

Creeping plant with
ovate leaves

Arthraxon

Erect plant with
linear leaves

201

201.

Fertile spikelet ±cylindrical
in

cross-section; callus pungent

Heteropogon

Fertile spikelet
±compressed; callus not

pungent

Dichanthium

202.

Spikelets laterally
compressed

Chrysopogon

Spikelets dorsally
compressed or

±cylindrical

203

203.

Pedicelled spikelet
reduced to pedicel;

panicle axes not terminating in spikelets

*Sorghastrum

Pedicelled spikelet
developed, rarely

reduced or suppressed but then panicle axes terminating in spikelets

204

204.

Pedicelled spikelets
reduced to narrow,

linear glumes; culm nodes glabrous or pubescent, not bearded

Vacoparis

Pedicelled spikelets
usually well

developed, rarely greatly reduced or absent, but then culm nodes bearded with

stiff hairs

205

205.

Lemma awns of sessile
spikelet absent or

1-1.5cm long if present

Sorghum

Lemma awns of sessile
spikelet 2-9cm

long

Sarga

206.

Inflorescence a spike
or spike-like raceme

or digitate pair of racemes; glumes as long as or longer than the florets

207

Inflorescence
otherwise or if a spike or

raceme lower glume shorter than the florets

212

207.

Inflorescence of
paired racemes (with

rarely a third raceme)

Dimeria

Inflorescence a single
spike or raceme

208

208.

Inflorescence a spike

Thaumastochloa

Inflorescence single
raceme

209

209.

Spikelets with
distinct bristles from

confluent tubercles

Thyridolepis

Spikelets with
indistinct bristles from

single tubercles, sometimes densely hairy

210

210.

211.

Plant shortly
rhizomatous

Neurachne

Plant stoloniferous

211

Annuals

Thedachloa

Perennials

Paraneurachne

212.

Spikelets with awns
arising from between

apical lobes of upper glume and lower lemma

*Melinis

Spikelets awnless or
with awn arising

terminally from lemmas or glumes

213

213

Spikelets in bunches
subtended by

bracts

Xerochloa

Spikelets not in
bunches subtended by

bracts

214

214.

Terminal spikelets
opening normally;

axillary spikelets cleistogamous

215

All spikelets opening
normally

217

215.


216.

Plants creeping

Calyptochloa

Plants tufted and wiry

216


Cleistogamous spikelets
in the axils of some leaf sheaths

Cleistochloa

Cleistogamous spikelets
produced from a leafy axillary shoot

Dimorphochloa

217.

Fertile lemma densely
silky hairy

Entolasia

Fertile lemma not or
indistinctly

hairy

218

218.

Inflorescence with
broad, flattened axis

with spikelets on one or both sides

219

Inflorescence
otherwise

220

219.

Spikelets closely
spaced on one side of

inflorescence axis

Stenotaphrum

Spikelets in distinct
clusters on both

sides of inflorescence axis

Uranthoecium

220.

Inflorescence branches
ending in

bristle-like appendages or spikelets subtended by bristles

221

Inflorescence branches
ending in spikelets

which are not subtended by bristles

227

221.

Spikelets falling at
maturity without any

bristles or spines attached

222

Spikelets falling at
maturity with bristles

or spines attached

225

222.

Spikelets subtended by
solitary bristles or

an involucre of bristles

Setaria

Spikelets in racemes
terminating in

bristle-like appendages

223

223.

Lower palea with wings
clasping margin of

upper floret

Holcolemma

Lower palea not
winged, not clasping upper

floret

224

224.

Upper lemma covered
with wrinkles, as long

as or longer than upper glume

Paspalidium

Upper lemma smooth,
shorter than upper

glume

Pseudoraphis

225.

Spikelets completely
surrounded by a ring

of spines or bristles

Cenchrus

Spikelets not
surrounded by spines or

bristles

226

226.

Lower glume up to 1/3
spikelet length

Chamaeraphis

Lower glume more than
1/3 spikelet

length

Paractaenum

227

Upper glume and lower lemma
with hooked

hairs

Ancistrachne

Upper glume and lower
lemma without hooked

hairs

228

228.


229.

Spikelet with a
bead-like swelling at

base

Eriochloa

Spikelet without a
bead-like swelling at

base

229

Upper lemma with a
shallow excavation and tuft of wool at the tip

*Lasiacis

Upper lemma rarely
excavated and then not woolly

230

230.

Fertile lemma less
firm or similar in

texture to glumes

231

Fertile lemma
decidedly firmer than the

glumes

238

231.

Lower glume 2/3
spikelet length

232

Lower glume to 1/2
spikelet length or

absent

233

232.

Inflorescence branches
stiff and spreading;

palea notched with a central depression

Walwhalleya

Inflorescence
consisting of only a few

spikelets, with branches not stiff and spreading; palea apex not notched

Alexfloydia

233.

Upper glume mucronate
to shortly awned

Alloteropsis

Upper glume obtuse to
acuminate, not awned

or mucronate

234

234.

Palea of upper floret
not completely

clasped by lemma

235

Palea of upper floret
completely clasped by

lemma

236

235.

Spikelet laterally
compressed

Dallwatsonia

Spikelet dorsally
compressed

Hymenachne

236.

Lower glume minute or
absent; spikelet

dorsally compressed

Digitaria

Lower glume c. 1/3
spikelet length;

spikelet terete or ±laterally compressed

237

237.

Lower glume minute or
absent; spikelet

dorsally compressed

Digitaria

Spikelet terete or ±
laterally

compressed

Steinchisma

238.

Rachilla prominent
between glumes and

between florets

239

Rachilla not prominent
between glumes and

between florets; fertile florets without appendage at base

242

239.

Rachilla swollen
beneath fertile florets

and usually with lateral appendages

240

Rachilla not swollen
beneath fertile

florets and without lateral appendages

241

240.

Appendages fused to
base of fertile floret;

spikelets laterally compressed, shortly pedicelled

Ichnanthus

Appendages, when
present, free from base of

fertile floret or absent; spikelets dorsally compressed, long pedicelled

Yakirra

241.

Articulations present
at the bases of spikelets, pedicels and inflorescence branches; upper glume 11-nerved

Arthragrostis

Articulations absent
on pedicels and inflorescence branches; upper glume 5-7-nerved

Whiteochloa

242.

Inflorescence an open
or contracted panicle branched more than once; spikelets not grouped along one side of ultimate

branches

243

Inflorescence once
branched, the branches digitate or borne on elongated axis; spikelets grouped along one side of branches

250

243.

Glumes slightly unequal
in length

244

Glumes very unequal in
length

246

244

Spikelets 4.5-6mm
long, borne at the ends of long branches

Homopholis

Spikelets 0.2-2 mm
long, on short branches

245

245.

Upper glume 3-nerved;
spikelets falling with glumes and compressed laterally

Cyrtococcum

Upper glume
5-9-nerved; spikelets disarticulating above glumes and compressed dorsally

Isachne

246.

Inflorescence a
spike-like panicle

Sacciolepis

Inflorescence an open
panicle

247

247.

248.


249.

Upper glumes somewhat
shorter than spikelet

Ottochloa

Upper glume as long as
spikelet

248

Fertile floret smooth
and shiny

Panicum

Fertile floret
transversely rugose

249

Culm base bulbous

*Zuloagaea

Culm base not bulbous

*Megathyrsus

250.

Lower glume absent or
represented by a minute scale

251

Lower glume present

252

251.

Lower lemma adjacent
to the inflorescence axis

*Axonopus

Lower lemma positioned
away from the inflorescence axis

Paspalum

252.

Spikelets awnless or
mucronate

253

Spikelets distinctly
awned

254

253.

Upper lemma rugose and
verrucose

Moorochloa

Upper lemma smooth

Urochloa

254.

Glumes more or less
equal and similar; decumbent grass of shaded habitats

Oplismenus

Glumes very unequal
and dissimilar; erect grass of wet habitats

Echinochloa

Scratchpads developed and conceived by (alphabetical): Ed Baker, Katherine Bouton Alice Heaton Dimitris Koureas, Laurence Livermore, Dave Roberts, Simon Rycroft, Ben Scott, Vince Smith