1.
2.
3.
|
Culms woody; leaves pseudopetiolate |
2 |
Culms usually herbaceous; if woody the leaves not pseudopetiolate |
5 |
|
Culms forming climbing vines |
||
Culms not forming climbing vines |
3 |
|
Auricles with bristles to 10 mm long |
||
Auricles without bristles |
4 |
|
4. |
Culms cylindrical in cross section |
|
Culms with an internodal groove on alternate sides |
||
5. |
Leaves spirally arranged |
|
Leaves arranged in two rows |
6 |
|
6. |
Spikelets morphologically or functionally unisexual |
7 |
Spikelets bisexual |
20 |
|
7. |
Spikelets with 1 floret, or more than 2 florets |
8 |
Spikelets with 2 florets |
12 |
|
8. |
Spikelets with 1 floret; male and female spikelets on same plant |
9 |
Spikelets with more than 2 florets; male and female spikelets on different plants |
10 |
|
9. |
Lemma asymmetrical with a ±lateral orifice |
|
Lemma pyriform with a terminal orifice |
||
10.
11. |
Short grasses with a compact raceme |
|
Tall reed-like grasses with a plumose panicle |
11 |
|
Leaf blades with only the midrib prominent and strongly sclerified |
||
Leaf blades with several wide strongly sclerified veins (including midrib) |
||
12. |
Upper lemma membranous, thinner than glumes |
13 |
Upper lemma hardened, thicker than glumes |
16 |
|
13. |
Male and female spikelets in separate inflorescences |
*Zea |
Male and female spikelets in different parts of the same inflorescence |
14 |
|
14.
15.
|
Inflorescence consisting of a female spikelet enclosed in a rigid shell, from which a raceme of male spikelets emerges apically |
|
Inflorescence spike-like with female spikelets basal and male apical |
15 |
|
Internode broader than and ± enclosing female spikelet |
||
Internode narrower than and ± enclosed by female spikelet |
||
16. |
Plants monoecioeus, i.e. with male and female spikelets on the same plant |
17 |
Plants dioecious i.e. with male and female spikelets on different plants |
18 |
|
17. |
Inflorescence a simple raceme, with 1-2 female spikelets at the base, the upper spikelets male |
|
Inflorescence a spike-like panicle with racemes of female spikelets below and solitary male spikelets above |
||
18. |
Inflorescence a contracted, spike-like panicle, not spatheate |
|
Inflorescences clustered together in dense, umbel-like spatheate heads |
19 |
|
19. |
Female heads with long stiff spines; inhabits coastal dunes |
|
Female inflorescences bracteate but not spiny; inhabits desert sand dunes |
||
20. |
Spikelets with 1 to many florets, if 2-flowered both florets or lower one bisexual |
21 |
Spikelets 2-flowered, lower floret male or barren, upper bisexual or female |
167 |
|
21. |
Inflorescence a solitary bilateral spike or raceme; spikelets usually with the broad side facing the axis |
22 |
Inflorescence a panicle or a unilateral or multilateral spike or raceme, rarely a bilateral spike but then the spikelets edgewise on to the axis |
38 |
|
22. |
Lemmas 3-5-nerved, hyaline to membranous |
23 |
Lemmas 5-10-nerved, hardened at maturity |
26 |
|
23. |
Lemmas awned |
|
Lemmas not awned |
24 |
|
24. |
Spikelets compressed laterally |
|
Spikelets compressed dorsiventrally |
25 |
|
25. |
Lemmas with lateral nerves extending to the apex |
|
Lemmas with very short lateral nerves |
||
26. |
Rachilla terminated by a fertile floret |
|
Rachilla prolonged beyond the uppermost fertile floret |
27 |
|
27. |
Spikelets in groups of more than 2 at each rachis node |
28 |
Spikelets solitary at each rachis node |
26 |
|
28. |
Spikelet 4-5-flowered, in groups of 2-4 (-7) |
|
Spikelet 1-flowered, in groups of 2 or 3 |
30 |
|
29. |
Spikelets in pairs at each rachis node |
|
Spikelets in 3s at each rachis node |
||
30. |
Annuals |
31 |
Perennials |
34 |
|
31. |
Glumes not keeled |
|
Glumes strongly keeled |
32 |
|
32. |
Glumes subulate |
|
Glumes not subulate |
33 |
|
33. |
Lemmas with nerves converging upwards |
|
Lemmas with nerves not converging upwards |
||
34. |
Rachis fragile, smooth |
|
Rachis tough, scabrous |
35 |
|
35. |
Plant with long creeping rhizomes |
|
Plant tufted, or with short rhizomes |
36 |
|
36 |
Leaf blades ± flat; glumes ± scabrous on nerves |
|
Leaf blades usually convolute or involute; glumes smooth |
37 |
|
37. |
Rachis internodes concave; glumes subacute to pointed |
|
Rachis internodes nearly flat; glumes truncate |
||
38. |
Fertile lemmas deeply or distinctly lobed |
39 |
Fertile lemmas entire, dentate or slightly lobed |
42 |
|
39. |
Spikelets 1-flowered |
|
Spikelets 2-or more-flowered, rarely 1-flowered |
40 |
|
40. |
Lemmas 9-lobed |
|
Lemmas 3-lobed |
41 |
|
41. |
Inflorescence a 1-sided raceme |
|
Inflorescence a panicle or spike |
||
42. |
Leaf blades hard, woody, needle-like |
|
Leaf blades not hard, woody, needle-like |
43 |
|
43. |
Leaf blades with transverse veins between main veins |
44 |
Leaf blades without transverse veins |
45 |
|
44. |
Glumes more or less equal; upper glume 7-nerved; lemmas awned |
|
Glumes very unequal; upper glume 3-5-nerved; lemmas mucronate |
||
45. |
Tall reed-like plants |
46 |
Plants not tall and reed-like |
47 |
|
46. |
Lemmas hairy; rachilla not hairy |
|
Lemmas not hairy; rachilla hairy |
||
47. |
Spikelets with 2 or more bisexual florets or if only one, with rachilla extension or sterile florets (empty lemmas) above it |
48 |
Spikelets with 1 bisexual floret, with or without male or sterile florets (empty lemmas) below it |
129 |
|
48. |
Spikelets with 3 or more florets or with 1or 2 florets and a rachilla extended beyond the upper floret |
49 |
Spikelets with 2 florets, without a rachilla extension |
122 |
|
49. |
Inflorescence racemose or spicate, or digitate or non-digitate |
50 |
Inflorescence a panicle |
76 |
|
50. |
Inflorescence digitate or subdigitate |
51 |
Inflorescence racemose or spicate |
62 |
|
51 |
Spikelet with at least 3 fertile florets |
52 |
Spikelet with 1 (rarely 2) fertile florets |
55 |
|
52. |
Axis of each inflorescence branch ending in a bristle; spikelet-bearing axis disarticulating |
|
Axis of each inflorescence branch ending in a spikelet; spikelet-bearing axis not disarticulating |
53 |
|
53. |
Spikelets 2-5; lemma 5-7-nerved; leaves mostly basal, setaceous |
|
Spikelets many; lemma 3-nerved; leaves not basally aggregated or setaceous |
54 |
|
54. |
Lemmas mucronate to shortly awned; upper glume 1-nerved |
|
Lemmas muticous; upper glumes 3-7-nerved |
||
55. |
Spikelet with 1 floret; lemmas unawned |
56 |
Spikelet with 2 or more florets; lemmas awned |
57 |
|
56. |
Glumes shorter than floret |
|
Glumes longer than floret |
||
57. |
Lowest lemma dorsally compressed (lying on front or back when placed on a flat surface) |
58 |
Lowest lemma laterally compressed (lying on the side when placed on a flat surface) |
59 |
|
58. |
Glumes unequal, shorter than florets |
|
Glumes subequal, longer than florets |
||
59. |
Inflorescence with more than 40 branches |
|
Inflorescence with less than 20 branches |
60 |
|
60. |
Awn of lowest lemma less than 2 mm long |
|
Awn of lowest lemma more than 3 mm long |
61 |
|
61. |
Lemmas very broad, wing-like |
|
Lemmas not very broad, not wing-like |
||
62. |
Spikelets unilateral (borne on one side of the inflorescence axis) |
63 |
Spikelets bilateral or multilateral (borne on opposite sides or all round the inflorescence axis) |
70 |
|
63. |
Spikelet with many fertile florets |
64 |
Spikelet with one fertile floret |
66 |
|
64. |
Lower glume 5-nerved; lemma 7-nerved |
|
Lower glume 1-nerved; lemma 1-3-nerved |
65 |
|
65. |
Lemma mucronate to shortly awned; spikelet rachilla and callus not hairy |
|
Lemma awnless; spikelet rachilla and callus hairy |
||
66. |
One glume only developed |
67 |
Both glumes well developed |
68 |
|
67. |
Spikelet with one floret; lemma 3-nerved |
|
Spikelet with many florets; lemma 3-nerved |
||
68. |
Spikelets falling entire; spikelets 6-18 mm long |
|
Spikelets fragmenting above glumes; spikelets less than 5.5 mm long |
69 |
|
69. |
Spikelets shortly pedicellate |
|
Spikelets sessile |
||
70. |
Spikelets disarticulating above glumes; rachilla extending beyond upper floret |
71 |
Spikelets falling with glumes; rachilla terminated by a floret |
73 |
|
71. |
Spikelets sessile, partially embedded in the rachis |
|
Spikelets pedicelled, not embedded in the rachis |
72 |
|
72. |
Lemmas awned; glumes not keeled |
|
Lemmas awnless; glumes keeled |
||
73. |
Inflorescence a true spike; spikelets sessile, sunken in hollows along the axis |
|
Inflorescence a spike-like raceme or spike-like panicle; spikelets at least shortly pedicellate and not sunken in hollows |
74 |
|
74. |
Spikelets burr-like with hooked spines |
|
Spikelets not burr-like |
75 |
|
75. |
Glume 1 |
|
Glumes 2 |
||
76. |
Glumes usually shorter than lowest floret and with the upper florets distinctly exserted; lemmas awnless or with a straight or curved awn from an entire or bifid apex, or several-awned or lobed |
77 |
Glumes (or the longer one) at least as long as or longer than the lowest floret, often as long as the spikelet and enclosing the florets; lemmas awnless or more often awned from the back or from the sinus of a 2-lobed tip, the awn usually geniculate, rarely the awn straight or curved from the tip |
103 |
|
77. |
Lemmas 1-3-nerved |
78 |
Lemmas 5-many nerved |
87 |
|
78. |
Lemmas 3-awned |
|
Lemmas 1-awned or without awns |
79 |
|
79. |
Lemmas notched, toothed or lobed |
80 |
Lemmas entire |
82 |
|
80. |
Glumes longer than lemmas, awned |
|
Glumes shorter than or equal to lemmas, not awned |
81 |
|
81. |
Spikelets borne in leaf-sheath axils |
|
Spikelets not borne in leaf-sheath axils |
||
82. |
Lemmas awned |
83 |
Lemmas not awned |
84 |
|
83. |
Spikelets in globose clusters; palea winged |
|
Spikelets in dense to loose panicles; palea not winged |
||
84. |
Lemmas with winged keels |
|
Lemmas without winged keels |
85 |
|
85. |
Ligule membranous |
|
Ligule a fringe of hairs |
86 |
|
86. |
Lemmas 1-nerved |
|
Lemmas 3-nerved |
||
87. |
Ovary hairy at apex |
88 |
Ovary glabrous at apex |
90 |
|
88. |
Leaf sheath margins joined |
|
Leaf sheath margins free |
89 |
|
89. |
Leaf blades 7-24 mm wide; lemmas bilobed |
|
Leaf blades 2-3 mm wide; lemmas 3-lobed |
||
90. |
Spikelets dimorphous, with bisexual spikelets surrounded by sterile ones |
91 |
Spikelets not dimorphous |
92 |
|
91. |
Spikelets disarticulating above the glumes; spikelet-bearing axes persistent |
|
Spikelets disarticulating with the glumes; spikelet bearing axes disarticulating |
||
92. |
Inflorescence a contracted panicle or a panicle of racemes |
93 |
Inflorescence an open panicle |
97 |
|
93. |
Glumes more or less equal |
94 |
Glumes very unequal |
96 |
|
94. |
Glumes not keeled |
|
Glumes keeled |
95 |
|
95. |
Leaf blades up to 3 mm long and basal, if present |
|
Leaf blades well developed and not mainly basal |
||
96. |
Glumes very dissimilar; lemmas awned |
|
Glumes similar; lemmas awnless |
||
97. |
Spikelets almost as wide as long |
|
Spikelets up to half as wide as long |
98 |
|
98. |
Lemmas ± rounded or flattened on back, at least below |
99 |
Lemmas keeled throughout |
100 |
|
99. |
Lemmas 7-9-nerved; leaf |
|
Lemmas 5-nerved; leaf |
||
100. |
Leaves not basally |
101 |
Leaves mostly basal; |
102 |
|
101. |
Lemmas keeled; palea |
|
Lemmas not keeled; |
||
102 |
Glumes as long the |
|
Glumes much shorter |
||
103. |
Ligule membranous |
104 |
Ligule a fringe of |
113 |
|
104. |
Lemmas 3-nerved |
|
Lemmas 5-9-nerved |
105 |
|
105. |
Lower floret male |
|
Lower floret bisexual |
106 |
|
106 |
Glumes more than 14 mm |
|
Glumes less than 10 mm |
107 |
|
107. |
Annuals |
108 |
Perennials |
110 |
|
108.
109. |
Panicle spicate |
|
Panicle open |
109 |
|
Awn witha ring of |
||
Awn without a ring of |
*Aira |
|
110. |
Lemmas 4-pointed, or |
111 |
Lemmas entire, |
112 |
|
111. |
Glumes long relative |
|
Glumes shorter than |
||
112 |
Lemma awn 4-9 mm long, |
|
Lemma awnless or with |
||
113. |
Leaf sheaths of the |
114 |
Leaf sheaths of the |
115 |
|
114. |
Spikelets with 3-8 |
|
Spikelets with 1 |
||
115. |
Lemmas awnless, |
116 |
Lemmas distinctly |
120 |
|
116. |
Lemmas entire, if |
|
Lemmas bilobed; if |
117 |
|
117. |
Grain not |
|
Grain longitudinally |
118 |
|
118. |
Lemma only hairy on |
|
Lemma ± evenly |
119 |
|
119. |
Lemma ± evenly |
|
Lemma with hairs over |
||
120 |
Hilum ± 1/2 length of |
|
Hilum up to 1/3 length |
121 |
|
121. |
Lemma ± evenly |
|
Lemma with hairs over |
122 |
|
122 |
Hairs on the lemma in |
|
Hairs on the lemma |
123 |
|
123. |
Lemmas either |
124 |
Lemmas neither |
128 |
|
124. |
Upper glume twice |
|
Glumes ± equal in |
125 |
|
125. |
Glumes (3-)5-11-nerved |
|
Glumes 1-3-nerved |
126 |
|
126. |
Bisexual florets 1 per |
|
Bisexual florets 2 per |
127 |
|
127. |
Lemma awn dorsal |
|
Lemma awn terminal |
||
128. |
Both lemmas |
|
Lower lemma hardened |
||
129. |
Glumes absent or |
130 |
Glumes well developed |
132 |
|
130. |
Spikelets 1-flowered |
|
Spikelets 3-flowered, |
131 |
|
131. |
Lemma and palea |
|
Lemma and palea |
||
132. |
Spikelets with 3 |
133 |
Spikelets with 1 |
137 |
|
133. |
Both sterile lemma |
|
Neither sterile lemmas |
134 |
|
134. |
Basal florets male |
|
Basal florets sterile |
135 |
|
135. |
Glumes minute |
|
Glumes relatively |
136 |
|
136. |
Sterile lemmas shorter |
|
Sterile lemmas longer |
||
137. |
Lemmas terete or |
138 |
Lemmas laterally |
145 |
|
138.
139.
|
Lemma with a |
139 |
Lemma with a single |
140 |
|
Awns glabrous |
||
Awns plumose |
||
140. |
Awn ± geniculate; |
141 |
Awn ± straight; |
144 |
|
141. |
Lemma margins usually |
142 |
Lemma margins not |
143 |
|
142. |
Hummock-forming |
|
Annuals or perennials, |
||
143. |
Lemma apex and lower |
|
Lemma apex glabrous or |
||
144. |
Lemma smooth, with a |
|
Lemma strongly |
||
145. |
Lemmas coriaceous to |
146 |
Lemmas membranous to |
151 |
|
146 |
Lemma awns |
147 |
Lemma awns geniculate |
148 |
|
147. |
Lemma with a |
|
Lemma with a single |
||
148. |
Lemma 1 awned and with |
149 |
Lemma 3-5 awned and |
150 |
|
149. |
Awn at least 15 mm |
|
Awn less than 10 mm |
||
150. |
Lemma with 5 awns |
|
Lemma with 3 awns |
||
151. |
Glumes similar in |
152 |
Glumes firmer in |
153 |
|
152. |
Panicle spike-like, |
|
Panicle open to |
||
153. |
Spikelets falling |
154 |
Spikelets |
156 |
|
154. |
Glumes blunt, ±united |
|
Glumes acute or awned, |
155 |
|
155 |
Lemma 3-nerved |
|
Lemma 5-nerved |
||
156. |
Glumes swollen at base |
|
Glumes not swollen at |
157 |
|
157. |
Glumes silky-woolly, |
|
Glumes not |
158 |
|
158. |
Spikelets at least 9mm |
|
Spikelets up to 7mm |
159 |
|
159. |
Glumes terminating |
160 |
Glumes obtuse or |
161 |
|
160. |
Lemmas awned; |
|
Lemmas not awned; |
||
161. |
Glumes with regularly |
|
Glumes smooth to rough |
162 |
|
162.
163.
|
Lemma awns more than |
163 |
Lemma awns, if |
164 |
|
Annuals |
||
Perennials |
||
164. |
Lemmas decidedly |
|
Lemmas less firm than |
165 |
|
165. |
Glumes at least 5.5mm |
|
Glumes less than 5mm |
166 |
|
166. |
Inflorescence axis |
|
Inflorescence often |
||
167. |
Spikelets breaking up |
168 |
Spikelets falling |
169 |
|
168. |
Glumes very unequal; |
|
Glumes ±equal; |
||
169. 170. |
Spikelets often |
170 |
Spikelets all alike |
206 |
|
Spikelets similar in shape and size, development |
171 |
|
Spiklets different in |
179 |
|
|
||
171. |
Paired spikelets |
172 |
Paired spikelets with |
174 |
|
172. |
Spikelets awnless |
|
Spikelets awned from |
173 |
|
173. |
Inflorescence a |
|
Inflorescence of |
||
174. |
Inflorescence a much |
|
Inflorescence a |
175 |
|
175. |
Spikelets in 3s, with |
|
Spikelets in pairs (3s |
176 |
|
176. |
Sessile spikelets |
|
Sessile and pedicelled |
177 |
|
177. |
Lower glume convex on |
|
Lower glume flat or |
178 |
|
178. |
Tufted grass with |
|
Creeping grass with no |
||
179. |
Rachis joints and |
180 |
Rachis joints and |
188 |
|
180. |
Sessile spikelet awned |
181 |
Sessile spikelet |
183 |
|
181. |
Pedicelled spikelets |
|
Pedicelled spikelets |
182 |
|
182. |
Pedicels ± flat |
|
Pedicels swollen |
||
183. |
Lower glume of sessile |
|
Lower glume of sessile |
184 |
|
184. |
Spikelets in opposite |
|
Spikelets on one side |
185 |
|
185. |
Spikelets of each pair |
|
Spikelets of each pair |
186 |
|
186. |
Lower floret of |
|
Lower floret of |
187 |
|
187. |
Lower glume of sessile |
|
Lower glume of sessile |
||
188. |
Inflorescence with |
189 |
Inflorescences |
197 |
|
189. |
Raceme 1 jointed, |
190 |
Racemes with several |
191 |
|
190. |
Racemes in tight |
|
Racemes loose and |
||
191. |
Fertile spikelets |
192 |
Fertile spikelets |
195 |
|
192. |
Lower part of awn as |
|
Lower part of awn |
193 |
|
193. |
Raceme bases terete |
|
Raceme bases flattened |
194 |
|
194. |
Racemes paired; leaves |
|
Racemes solitary; |
||
195. |
Fertile spikelets not |
|
Fertile spikelets |
196 |
|
196. |
Fertile spikelets |
|
Fertile spikelets |
||
197. |
Rachis joints and |
198 |
Rachis joints and |
199 |
|
198. |
Inflorescence more |
|
Inflorescence an |
||
199. |
Inflorescence of |
200 |
Inflorescence a |
202 |
|
200. |
Creeping plant with |
|
Erect plant with |
201 |
|
201. |
Fertile spikelet ±cylindrical |
|
Fertile spikelet |
||
202. |
Spikelets laterally |
|
Spikelets dorsally |
203 |
|
203. |
Pedicelled spikelet |
|
Pedicelled spikelet |
204 |
|
204. |
Pedicelled spikelets |
|
Pedicelled spikelets |
205 |
|
205. |
Lemma awns of sessile |
|
Lemma awns of sessile |
||
206. |
Inflorescence a spike |
207 |
Inflorescence |
212 |
|
207. |
Inflorescence of |
|
Inflorescence a single |
208 |
|
208. |
Inflorescence a spike |
|
Inflorescence single |
209 |
|
209. |
Spikelets with |
|
Spikelets with |
210 |
|
210.
211. |
Plant shortly |
|
Plant stoloniferous |
211 |
|
Annuals |
||
Perennials |
||
212. |
Spikelets with awns |
|
Spikelets awnless or |
213 |
|
213 |
Spikelets in bunches |
|
Spikelets not in |
214 |
|
214. |
Terminal spikelets |
215 |
All spikelets opening |
217 |
|
215. 216. |
Plants creeping |
|
Plants tufted and wiry |
216 |
|
|
||
Cleistogamous spikelets |
||
Cleistogamous spikelets |
||
217. |
Fertile lemma densely |
|
Fertile lemma not or |
218 |
|
218. |
Inflorescence with |
219 |
Inflorescence |
220 |
|
219. |
Spikelets closely |
|
Spikelets in distinct |
||
220. |
Inflorescence branches |
221 |
Inflorescence branches |
227 |
|
221. |
Spikelets falling at |
222 |
Spikelets falling at |
225 |
|
222. |
Spikelets subtended by |
|
Spikelets in racemes |
223 |
|
223. |
Lower palea with wings |
|
Lower palea not |
224 |
|
224. |
Upper lemma covered |
|
Upper lemma smooth, |
||
225. |
Spikelets completely |
|
Spikelets not |
226 |
|
226. |
Lower glume up to 1/3 |
|
Lower glume more than |
||
227 |
Upper glume and lower lemma |
|
Upper glume and lower |
228 |
|
228. 229. |
Spikelet with a |
|
Spikelet without a |
229 |
|
Upper lemma with a |
||
Upper lemma rarely |
230 |
|
230. |
Fertile lemma less |
231 |
Fertile lemma |
238 |
|
231. |
Lower glume 2/3 |
232 |
Lower glume to 1/2 |
233 |
|
232. |
Inflorescence branches |
|
Inflorescence |
||
233. |
Upper glume mucronate |
|
Upper glume obtuse to |
234 |
|
234. |
Palea of upper floret |
235 |
Palea of upper floret |
236 |
|
235. |
Spikelet laterally |
|
Spikelet dorsally |
||
236. |
Lower glume minute or |
|
Lower glume c. 1/3 |
237 |
|
237. |
Lower glume minute or |
|
Spikelet terete or ± |
||
238. |
Rachilla prominent |
239 |
Rachilla not prominent |
242 |
|
239. |
Rachilla swollen |
240 |
Rachilla not swollen |
241 |
|
240. |
Appendages fused to |
|
Appendages, when |
||
241. |
Articulations present |
|
Articulations absent |
||
242. |
Inflorescence an open |
243 |
Inflorescence once |
250 |
|
243. |
Glumes slightly unequal |
244 |
Glumes very unequal in |
246 |
|
244 |
Spikelets 4.5-6mm |
|
Spikelets 0.2-2 mm |
245 |
|
245. |
Upper glume 3-nerved; |
|
Upper glume |
||
246. |
Inflorescence a |
|
Inflorescence an open |
247 |
|
247.
248. 249. |
Upper glumes somewhat |
|
Upper glume as long as |
248 |
|
Fertile floret smooth |
||
Fertile floret |
249 |
|
Culm base bulbous |
*Zuloagaea |
|
Culm base not bulbous |
||
250. |
Lower glume absent or |
251 |
Lower glume present |
252 |
|
251. |
Lower lemma adjacent |
|
Lower lemma positioned |
||
252. |
Spikelets awnless or |
253 |
Spikelets distinctly |
254 |
|
253. |
Upper lemma rugose and |
|
Upper lemma smooth |
||
254. |
Glumes more or less |
|
Glumes very unequal |